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In Chrome, and in old versions of Firefox (and Firefox after this bug is resolved), that causes a smooth animation because transform: none uses the identity transform.
In release versions of chrome and older versions of Firefox, .animation { transform: perspective(0) translateZ(0) } was a no-op in the above test-case, but after #6320perspective(0) is perspective(1px), and that's a fairly drastic behavior change. In fact, there's no way an author can get an identity transform into perspective() other than by not specifying transform at all, which seems unfortunate / wrong.